According to the ideology of the Darkness and Sword Logic, the Final Shape is a state where one singular species or individual has overcome all other life to become the last living things in existence. In the "Unveiling", we learned that it was the Pattern that became the Vex which embodied this ideology, which the Winnower admired. Within the game, it is the Hive who actively pursue the Final Shape through Sword Logic, and the Darkness has taken an interest in Guardians because it believes we can become the Final Shape.
The thing is, the existence of trillions of timelines and other universes adds an additional layer to things. For instance, a single race could become the last thing in existence in one timeline, but they may have become extinct in [i]another timeline[/i] where another race became the Final Shape, and so on. For instance, there could be a timeline where the Devil Splicers won against the Last City; timelines where Crota and Oryx never died; a timeline where Ghaul won the Red War and successfully held onto the Light he took.
Or perhaps even ourselves: a Dark Guardian who so embraced the Darkness they eventually became the Final Shape in their timeline/universe.
This is where the Vex come into the picture. The Vex extend their reach across not just one universe's past, present and future, but all possible timelines and all possible universes. They created Atheon in an attempt to create ontological weapons and write themselves into the fabric of reality. They created Panoptes and the Infinite Forest to find a future where their goals are achieved.
To be honest, a part of me has begun to wonder whether we merely destroy [i][b]our timeline's[/b][/i] versions of Atheon and Panoptes, or whether they were singular existences for the Vex Collective, but that's a topic for another time.
Because of the Vex's nature, I am rather certain that the future story Destiny is going towards will eventually extend towards those different universes and timelines, because the true scope of the Final Shape isn't simply to be the Final Shape in one universe, in one timeline. If the Vex and the Pattern in the field of possibility are any indications, the Final Shape needs to extend across all timelines and all universes.
In other words, any being, any race who became the Final Shape in their respective timeline or universe would inevitably turn their gaze towards other timelines, other universes, to eliminate any potential threats to them from those realities and timelines, or to ensure their own supremacy across timelines and realities, just as the Vex would.
Perhaps this could open up the possibility of a future villain in Destiny 2 being our own Dark Guardian counterpart, from one of the countless timelines where the Darkness won and our Guardian embraces the Darkness. After all, we don't really know what happened in those dark timelines Elsie has lived through after she's pulled back in time via the time-loop.
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I am the Final Shape........👻